Toby S. Daly-Engel

Assistant Professor | Ocean Engineering and Marine Sciences

Contact Information

Personal Overview

Dr. Toby’s research uses a combination of genomics, field ecology, and modeling to study mating systems, habitat use, and the impacts of climate change on shark populations.  She is interested in how reproductive behaviors, particularly from a female fitness perspective, interact with the environment to influence species, populations, individuals, and genomes. Dr. Toby (she/her) grew up in upstate New York and got her BA in Biology at Oberlin College. She worked in Boston for two years before going on to get her Master’s and PhD in Zoology with a specialization in Ecology, Evolution, and Conservation Biology at the University of Hawaii. She won accolades as an Assistant Professor at the University of West Florida, where she ran a shark tagging field program and helped found the West Florida chapter of the Association for Women in Science, before moving her research to the Florida Institute of Technology. As director of the Florida Tech Shark Conservation Lab, Dr. Toby and her students work with collaborators from state and federal agencies, non-governmental organizations, and public groups to study how reproduction and rising temperatures shape movement patterns in large migratory fishes, identify cryptic species, and work to increase representation for women in science, especially women of color.

Educational Background

Ph.D. Zoology, specializing in Ecology, Evolution, and Conservation Biology, University of Hawaii (2009)

M.S. Zoology, University of Hawaii (2007)

B.A. Biology, Oberlin College (2000)

Current Courses

BIO 1020_02, Advanced Biological Discovery (Honors)

MAR 3701, Evolution

MAR 4411/5031, Conservation Genetics

MAR 4100/5511, Elasmobiology (Biology of Sharks, Skates, and Rays)

Selected Publications

Drymon, J.M., K.E. Schweiss, E. Seubert, R.N. Lehman, T.S. Daly-Engel, M. Pfleger, and N.M. Phillips (2020) Swimming against the flow – testing the potential for environmental DNA to detect bull sharks (Carcharhinus leucas) across a dynamic deltaic interface. Ecology and Evolution,

Weber, H.K., C.M. Jones, M.J. Ajemian, M.K. McCallister, B.L. Winner, G.R. Poulakis, L.D. Hollensead, D. Zapf, J.D. Swenson, J.M. Hendon, T.S. Daly-Engel, and N.M. Phillips(2020) Genetic evidence supports a range extension of the Brazilian Cownose Ray, Rhinoptera brasiliensis, in the western North Atlantic. Environmental Biology of Fishes, 98(2), 577-582

Eble, J.A., T.S. Daly-Engel, J.D. DiBattista, A. Koziol, and M.R. Gaither (2020) Chapter 1. Marine Environmental DNA: Approaches, Applications, and Opportunities. In: Advances in Marine Biology 85: 1-23

Shiffman, D., M. Ajemian, J. Carrier, T.S. Daly-Engel, M. Davis, N. Dulvy, R.D. Grubbs, N.A. Hinojosa, J. Imhoff, M. Kolmann, C. Nash, M. Paig-Tran, E.E. Peele, R. Skubel, B. Wetherbee, L. Whitenack, and J. Wyffels (2020) Trends in Chondrichthyan research: An analysis of three decades of conference abstracts. Copeia 108(1), 122-131

Davis, M.M., P. Suárez-Moo, and T.S. Daly-Engel (2019) Genetic structure and congeneric range overlap among sharpnose sharks (genus Rhizoprionodon) in the Northwest Atlantic Ocean. Canadian Journal of Fisheries and Aquatic Sciences 76(7): 1203-1211

Rowley, A., L. Locatello, A. Kahrl, M. Rego, A. Boussard, E. Garza-Gisholt, R. Kempster, S.P. Collins, E. Giacomello, M.C. Follesa, C. Porcu, J. Evans, F. Hazin, F. Garcia-Gonzalez, T.S. Daly-Engel, C. Mazzoldi and J.LFitzpatrick (2019) Sexual selection and the evolution of sperm morphology in sharks. Journal of Evol. Biology, 32(10), 1027-1035

Rowley A., T.S. Daly-Engel, and J.L. Fitzpatrick (2018) Testes size increases with sperm competition risk and intensity in bony fish and sharks. Behavioral Ecology Dec. 27, 1–8. doi:10.1093/beheco/ary174

Daly-Engel, T.S., A. Koch, J.M. Anderson, C. Cotton, and R.D. Grubbs (2018) Description of a new deep-water dogfish shark from Hawaii, with comments on the Squalus mitsukurii species complex in the West Pacific. ZooKeys 798: 135-157 doi: 10.3897/zookeys.798.28375

Huveneers C., K. Apps, E.E. Becerril-García, B. Bruce, P.A. Butcher, A.B. Carlisle, T.K. Chapple, H.M. Christiansen, G. Cliff, T.H. Curtis, T.S. Daly-Engel, H. Dewar, M.L. Dicken, M.L. Domeier, C.A.J. Duffy, R. Ford, M.P. Francis, G.C.A. French, F. Galván-Magaña, E. García-Rodríguez, E. Gennari, B. Graham, B. Hayden, E.M. Hoyos-Padilla, N.E. Hussey, O.J.D. Jewell, S.J. Jorgensen, A.A. Kock, C.G. Lowe, K. Lyons, L. Meyer, G. Oelofse, E.C. Oñate-González, H. Oosthuizen, J.B. O’Sullivan, K. Ramm, G. Skomal, S. Sloan, M.J. Smale, O. Sosa-Nishizaki, E. Sperone, E. Tamburin, A.V. Towner, M.A. Wcisel, K.C. Weng, J.M. Werry (2018) Future Research Directions on the ‘Elusive’ White Shark. Frontiers in Marine Science 5: 455Davis, M.M, P. Suárez-Moo, and T.S. Daly-Engel (2018) Genetic structure and range expansion among sharpnose sharks (Genus Rhizoprionodon) in the Northwest Atlantic Ocean. Canadian Journal of Fisheries and Aquatic Sciences 10.1139/cjfas-2018-0019

Pfleger, M.O., R.D. Grubbs, C. Cotton, and T.S. Daly-Engel (2018) Squalus clarkae sp. nov., a new dogfish shark from the Northwest Atlantic and Gulf of Mexico, with comments on the Squalus mitsukurii species complex. Zootaxa  4444 (2): 101–119

Daly-Engel, T.S., I. Baremore, R.D. Grubbs, S. Gulak, R.T. Graham, and M. Enzenauer (2018) Resurrection of the sixgill shark Hexanchus vitulus Springer &Waller, 1969 (Hexanchiformes, Hexanchidae), with comments on its distribution in the northwest Atlantic Ocean. Marine Biodiversity:

Larson, S., T.S. Daly-Engel, and N. Phillips (2017) Ch. 3: Review of Current Conservation Genetic Analyses of Northeast Pacific Sharks. In Advances in Marine Biology: Northeast Pacific Shark Biology, Research and Conservation. Eds: S. Larson and D. Lowry. Vol. 77, Oxford: Academic Press, pp. 79-110

Selkoe, K.A., E.A. Treml, J.L.K. Wren, O. Gaggiotti, M. Donovan, K. Andrews, I. Baums, M.A. Bernal, C. Bird, H. Bolick, B. Bowen, R. Coleman, G.T. Concepcion, M.T. Craig, T.S. Daly-Engel, J.D. DiBattista, J. Eble, I. Fernandez-Silva, E. Franklin, A. Friedlander, M.R. Gaither, J. Gove, M. Iacchei, Y. Jia, M.A.J. Rivera, L.A. Rocha, J. Reece, D. Skillings, S. Santos, Z. Szabo, M. Timmers, L. Wedding, G.J. Williams, N. Whitney, and R.J. Toonen (2016) Community genetics of Hawaiian reefs reflect habitat, history, thermal stress and ecology. Proceedings of the Royal Society of London B 283: 20160354

Daly-Engel, T.S., R.L. Smith, D.S. Finn, M.E. Knoderbane, I.C. Phillipsen, and D.A. Lytle (2012a) 17 novel polymorphic microsatellite markers for the giant water bug, Abedus herberti (Belostomatidae). Conservation Genetics Resources 4: 979-981

Daly-Engel, T.S., K.M. Duncan, K.N. Holland, J.P. Coffey, H.A. Nance, R.J. Toonen, and B.W. Bowen (2012b) Global phylogeography with mixed-marker analysis reveals male-mediated dispersal in the endangered scalloped hammerhead shark (Sphyrna lewini). PLoS ONE 7(1): e29986

Fitzpatrick, J.L., R.M. Kempster, T.S. Daly-Engel, S.P. Collin, and J.P. Evans (2012) Assessing the potential for postcopulatory sexual selection in elasmobranchs. The Journal of Fish Biology's 2012 special issue, The Current Status of Elasmobranchs: Biology, Fisheries and Conservation. DOI: 10.1111/j.1095-8649.2012.03256.x

Daly-Engel, T.S., J.E. Randall, and B.W. Bowen (2012c) Is the Great Barracuda (Sphyraena barracuda) a reef fish or a pelagic fish? The phylogeographic perspective. Marine Biology 159: 975-985

Toonen, R.J., K.R. Andrews, I.B. Baums, C.E. Bird, G.T. Concepcion, T.S. Daly-Engel, J.A. Eble, A. Faucci, M.R. Gaither, M. Iacchei, J.B. Puritz, J.K. Schultz, D.J. Skillings, M.A. Timmers, and B.W. Bowen (2011) Defining boundaries for ecosystem-based management: a multispecies case study of marine connectivity across the Hawaiian Archipelago. Journal of Marine Biology 2011: 1-13

Cotton, C.F., R.D. Grubbs, T.S. Daly-Engel, P.D. Lynch, and J.A. Musick (2011) Age, growth and reproduction of Squalus cf. mitsukurii from Hawaiian waters. Marine and Freshwater Research 62: 811-822

Daly-Engel, T.S., R.D. Grubbs, K. Feldheim, B.W. Bowen, and R.J. Toonen (2010) Is multiple mating beneficial or unavoidable? Low multiple paternity and genetic diversity in the shortspine spurdog, Squalus mitsukurii. Marine Ecology Progress Series 403: 255-267

Ebert, D.A., W.T. White, K.J. Goldman, L.J.V. Compagno, T.S. Daly-Engel, and R.D. Ward (2010) Reevaluation and redescription of Squalus suckleyi (Girard, 1854) from the North Pacific, with comments on the Squalus acanthias subgroup (Squaliformes: Squalidae: Squalus). Zootaxa 2612: 22-40

Nance, H.A., T.S. Daly-Engel, and P.B. Marko (2009) New microsatellite loci for the endangered scalloped hammerhead shark, Sphyrna lewini. Molecular Ecology Resources 9: 955-957

Bienfang, P., B. Oben, S. DeFelice, P. Moeller, K. Huncik, P. Oben, R. Toonen, T. Daly-Engel, and B. Bowen (2008) Ciguatera: Detection of neurotoxins in carnivorous reef fish from the coast of Cameroon, West Africa. African Journal of Marine Science 30: 533-540

Daly-Engel, T.S., R.D. Grubbs, B.W. Bowen, and R.J. Toonen (2007) Frequency of multiple paternity in an unexploited tropical population of sandbar sharks, Carcharhinus plumbeus. Canadian Journal of Fisheries and Aquatic Sciences 64: 198-204

Daly-Engel, T.S., R.D. Grubbs, K.N. Holland, R.J. Toonen, and B.W. Bowen (2006) Assessment of multiple paternity in single litters from three species of carcharhinid sharks in Hawaii. Environmental Biology of Fishes 76: 419-424

Duncan, K.M. and T.S. Daly-Engel (2006) Using forensic science problems as teaching tools. The Science Teacher Nov. 06: 38-43

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